behavior/neurological
• from 2 weeks of age, mutant mice lean from one side to the other as they walk and frequently fall over sideways
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nervous system
• the outer marginal layer of the cortex is populated by numerous ectopic cells
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• neurons of the mutant hippocampal gyrus are scattered throughout several irregular layers
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• the cortical layers of mutant mice are scrambled
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• Purkinje and granule cells of the mutant cerebellum are scrambled
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• Purkinje and granule cells of the mutant cerebellum are scrambled
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